Toona hexandra

(Wall.) M.Roem. (1846)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Meliaceae Genus: Toona

australian red cedar

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Description

Key Characters:

Growth Form: Fast–growing trees to ca. 20–30 m tall.

Stems: Bark thick, fibrous, bitter, astringent.

Roots:

Leaves: Leaves compound (pinnate). Alternate. Leaflets narrowly ovate to lanceolate, usually 3–6 dm long, leaflets 10–14(–18), 4.5–16 cm long. Apex acute to caudate. Base inequilateral. Surfaces glabrous or at least sparsely pubescent. Leaflet margins entire. Petiolate. Stipules absent.

Flowers: Flowers in many-branched thyrses. Flowers unisexual (with well–developed rudiments of the opposite sex present). Calyx (4)5-lobed, lobes ca. 1 mm long. Corolla of 5 petals, distinct, white, ca. 5–6 mm long, margins ciliate, longer than calyx in bud, imbricate, adnate at base to a short androgynophore. Stamens 5, distinct, arising from the androgynophore, staminal filaments ca. 2 mm long, staminodes absent; anthers of staminate flowers yellow, dehiscent; on pistillate flowers smaller, brown and shriveled, indehiscent, not producing pollen. Ovary superior, 5-celled; ovules 6–10 per cell, ovary and style glabrous; style capitate; in staminate flowers ovary more slender, cells well–developed and style longer.

Fruit: Capsules reddish brown; pendulous; septicidal; with some lighter spots; ellipsoid; 2–2.8 cm long; opening by 5–valves; the columella 5–angled. Seeds brown; thin; dry; the embryo portion ca. 1–2 mm long; with 2 wings; the longer one ca. 5–12 mm long; the other ca. ½ as long.

Ploidy: 2n = 56

Habitat: Mesic to wet; disturbed habitats.

Elevation Range: 25–610 m.

Historical Distribution

Synonyms (66)

Uses and Culture

USES

Ethnobotanical Images

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents

Water/Ocean
Wind

Vegetative Reproduction

Pollinators

Specimens

Bibliography

Name Published In: Fam. Nat. Syn. Monogr. 1: 139 (1846)

Other References

Wagner et al. 1990:920 (K, O, L, M, H [as Toona ciliata var. australis]); Herbst & Wagner 1999:24 (NOM:Syn. T. ciliata var. australis); Staples & Herbst 2005:398 (KEY), 401 (DESCR); Oppenheimer 2007:27 (Mo)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Toona hexandra (Wall.) M.Roem. honolulu, uh campus Under a canopy of Sapindus, dry off gulch floor terrain. PRESERVED_SPECIMEN woolford, h.a. Oahu BISH 3/12/1952
2 Toona hexandra (Wall.) M.Roem. mokuleia forest reserve, 5 mi. s of haleiwa roadside PRESERVED_SPECIMEN little jr., e.l. collector number: 31044 Oahu BISH 8/11/1976
3 Toona hexandra (Wall.) M.Roem. honolulu; foster botanic garden Planted tree on the property. PRESERVED_SPECIMEN lau, j. collector number: 3003 Hawaii BISH 12/9/1986
4 Toona hexandra (Wall.) M.Roem. Hilo? PRESERVED_SPECIMEN Bryan, L.W. 2 Hawaii BISH 12/26/1956
5 Toona hexandra (Wall.) M.Roem. koloa district, mountains above kalaheo, along road between alexander reservoir and kanaele. PRESERVED_SPECIMEN t. w. flynn, m. koehler, s. walsh, d. wolkis, k. brock & j. bernard 8726 F 4/6/2017
6 Toona hexandra (Wall.) M.Roem. Wahiawa Mts. Along middle to lower half of road to relay towers, Mt. Kahili. PRESERVED_SPECIMEN Tim Flynn 583 Kauai PTBG 9/7/1983
7 Toona hexandra (Wall.) M.Roem. makua kea'au forest reserve, wai'anae range PRESERVED_SPECIMEN c. r. annable 3076 MU 10/24/1996
8 Toona hexandra (Wall.) M.Roem. Makua Kea'au Forest Reserve, Wai'anae Range Mesic forest PRESERVED_SPECIMEN C. R. Annable 3076 NY 10/24/1996
9 Toona hexandra (Wall.) M.Roem. Honolulu, Makiki Hgts., Tantalus Drive. PRESERVED_SPECIMEN W. M. Bush 612 Oahu PTBG 5/5/1977
10 Toona hexandra (Wall.) M.Roem. MAUI: W. Maui, Lahaina District, Honokahua. Degraded mesic forest, east side of valley. PRESERVED_SPECIMEN Hank Oppenheimer H70096 Maui PTBG 7/19/2000
11 Toona hexandra (Wall.) M.Roem. Wahiawa drainage, North of Hulua, south of Waimea- Koloa district boundary, Lihue- Koloa Forest reserve, north of Kanaele swamp, Metrosideros polymorpha- Dicranopteris linearis wet forest PRESERVED_SPECIMEN Steve Perlman 23476 Kauai PTBG 5/23/2013
12 Toona hexandra (Wall.) M.Roem. Upper Waiakea Forest along Tree Planting Road approached from Stainback Highway. Wet mesic forest. PRESERVED_SPECIMEN J. Regalado 641 Hawaii PTBG 5/10/1993
13 Toona hexandra (Wall.) M.Roem. Hana, E maui, along Hana Hwy., about 10 mi from Hana town PRESERVED_SPECIMEN Spence, G. 74 Maui US 10/15/1972
14 Toona hexandra (Wall.) M.Roem. Mountains above Kalaheo, along road between Alexander Reservoir and Kanaele. Disturbed mesic forest PRESERVED_SPECIMEN Tim Flynn 8726 Kauai PTBG 4/6/2017
15 Toona hexandra (Wall.) M.Roem. Wailua, Wailua Game Reserve. Vegetation on plots stripmined for bauxite ore in 1958-60. Tree plot area. Fertilizer applied topically. PRESERVED_SPECIMEN Richard A. Howard 20226 Kauai PTBG 10/20/1985
16 Toona hexandra (Wall.) M.Roem. Kalaupapa, east end of peninsula. Northing: 2343087.65 Easting: 712867.62 PRESERVED_SPECIMEN Hank Oppenheimer H60727 Molokai PTBG 6/30/2007
17 Toona hexandra (Wall.) M.Roem. Kaiholena Gulch. PRESERVED_SPECIMEN David Palumbo 118 Lanai PTBG 6/3/1983
18 Toona hexandra (Wall.) M.Roem. honolulu, wilhelmina rise PRESERVED_SPECIMEN d. mitchell s.n. Oahu US
19 Toona hexandra (Wall.) M.Roem. hana, east maui, along hana hwy., about 10 miles from hana town. PRESERVED_SPECIMEN g. spence 74 Maui US 10/15/1972
20 Toona hexandra (Wall.) M.Roem. E part of Hawaii, Waiakea Arboretum, Waiakea Forest Preserve, 5 mi S of Hilo PRESERVED_SPECIMEN Little Jr., E. L. 23041 Hawaii US 6/2/1968
21 Toona hexandra (Wall.) M.Roem. Hana, East Maui, along Hana Hwy., about 10 miles from Hana town. PRESERVED_SPECIMEN Spence, G. 74 Maui US 10/15/1972
22 Toona hexandra (Wall.) M.Roem. Honolulu, Wilhelmina Rise PRESERVED_SPECIMEN Mitchell, D. s.n. Oahu US 10/1/1943
23 Toona hexandra (Wall.) M.Roem. NW part of Oahu, Mokulei Forest Reserve, 5 mi S of Haleiwa PRESERVED_SPECIMEN Little Jr., E. L. 31044 Oahu US 8/11/1976
24 Toona hexandra (Wall.) M.Roem. Koloa District, mountains above Kalaheo, along road between Alexander Reservoir and Kanaele. Disturbed mesic forest PRESERVED_SPECIMEN Flynn, T. W.; Koehler, M. S.; Walsh, S. K.; Wolkis, D.; Brock, K.; Bernard, J. 8726 Kauai US 4/6/2017
25 Toona hexandra (Wall.) M.Roem. Vicinity of Maunahui. Northing: 2339129.18; Easting: 712863.23. PRESERVED_SPECIMEN Hank Oppenheimer H110620 Molokai PTBG 11/27/2006
26 Toona hexandra (Wall.) M.Roem. PRESERVED_SPECIMEN collector unknown usfpl 17997 Maui US
27 Toona hexandra (Wall.) M.Roem. Honolulu; Foster Botanic Garden In full sun. PRESERVED_SPECIMEN Lau, J. 3003 Oahu BISH 12/9/1986
28 Toona hexandra (Wall.) M.Roem. hana, e maui, along hana hwy., about 10 mi from hana town PRESERVED_SPECIMEN g. spence 74 Kahoolawe US 10/15/1972
29 Toona hexandra (Wall.) M.Roem. nw part of oahu, mokulei forest reserve, 5 mi s of haleiwa PRESERVED_SPECIMEN e. l. little jr. 31044 Oahu US 8/11/1976
30 Toona hexandra (Wall.) M.Roem. Hilo? PRESERVED_SPECIMEN Bryan, L. W. 2 Hawaii US 12/26/1956
31 Toona hexandra (Wall.) M.Roem. hilo? PRESERVED_SPECIMEN l. bryan 2 Hawaii US 12/26/1956
32 Toona hexandra (Wall.) M.Roem. Hawaii PRESERVED_SPECIMEN Adee, K.T. s.n. Hawaii BISH
33 Toona hexandra (Wall.) M.Roem. kamanele park PRESERVED_SPECIMEN ishikawa, s. collector number: 37 Oahu BISH 1/17/1972
34 Toona hexandra (Wall.) M.Roem. honolulu, tantalus PRESERVED_SPECIMEN weber, p.w. Oahu BISH 5/25/1945
35 Toona hexandra (Wall.) M.Roem. Along jeep road to Kanaele Swamp (Wahiawa Bog) between bog & Alexander Reservoir PRESERVED_SPECIMEN Wagner, W.L. 5001 Kauai BISH 8/27/1983
36 Toona hexandra (Wall.) M.Roem. Waiakea Arboretum,Waiakea Forest Preserve,5m,S.of Hilo Secondary forest. PRESERVED_SPECIMEN Little Jr., E.L. 23041 Hawaii BISH 6/2/1968
37 Toona hexandra (Wall.) M.Roem. honolulu harbor PRESERVED_SPECIMEN adee, k.t. collector number: s.n. Oahu BISH
38 Toona hexandra (Wall.) M.Roem. east maui, hana, along highway 10 mi. from town PRESERVED_SPECIMEN spence, g. collector number: 74 Molokai BISH 10/15/1972
39 Toona hexandra (Wall.) M.Roem. Waiahole, Water Reserve PRESERVED_SPECIMEN Saito, F. s.n. Oahu BISH 5/22/1962
40 Toona hexandra (Wall.) M.Roem. Koloa District, mountains above Kalaheo, along road between Alexander Reservoir and Kanaele. Disturbed mesic forest PRESERVED_SPECIMEN T. W. Flynn, M. Koehler, S. Walsh, D. Wolkis, K. Brock & J. Bernard 8726 Kauai US 4/6/2017
41 Toona hexandra (Wall.) M.Roem. Cultivated near Sinclair Library, University of Hawaii Manoa Campus PRESERVED_SPECIMEN Herbst, D.R. 636 Oahu BISH 10/7/1967
42 Toona hexandra (Wall.) M.Roem. Honolulu: Kamanele Park PRESERVED_SPECIMEN Degener, O. 8776 Oahu BISH 5/1/1932
43 Toona hexandra (Wall.) M.Roem. Honolulu, Kamanele Park PRESERVED_SPECIMEN Degener, O. 8738 Oahu BISH 4/15/1905
44 Toona hexandra (Wall.) M.Roem. Mokuleia Forest Reserve, 5 mi S of Haleiwa roadside PRESERVED_SPECIMEN Little Jr., E.L. 31044 Oahu BISH 8/11/1976
45 Toona hexandra (Wall.) M.Roem. along jeep road to kanaele swamp (wahiawa bog) between bog & alexander reservoir Vine spreading through trees. PRESERVED_SPECIMEN wagner, w.l. mill, s.w.; d.wier; g.wier collector number: 5001 Oahu BISH 8/27/1983
46 Toona hexandra (Wall.) M.Roem. vicinity of maunahui mesic slopes PRESERVED_SPECIMEN oppenheimer, h.l. perlman, s. collector number: h110620 Oahu BISH 11/27/2006
47 Toona hexandra (Wall.) M.Roem. Koloa District. Wahiawa drainage, North of Hulua, south of Waimea-Koloa district boundary, Lihue-Koloa Forest reserve, north of Kanele swamp Metrosideros polymorpha-Dicranopteris linearis wet forest. Antidesma platyphylla, Ilex anomala, Polyscias waialeale, Scaevola gaudichaudiana, Bobea brevipes, Psychotria mariniana, Cyanea kahilensis, weeds: Psidium cattleianum PRESERVED_SPECIMEN Perlman, S. 23476 Kauai BISH 5/23/2013
48 Toona hexandra (Wall.) M.Roem. Wailua Game Reserve, above Wailua. Begetaion on plots stripmined for bauxite ore in 1958-60. Tree plot area. Fertilizer applied topically PRESERVED_SPECIMEN Howard, R.A. 20226 Kauai BISH 10/20/1985
49 Toona hexandra (Wall.) M.Roem. Hawaii County, Hawaii Island, Roadside on Stainback Highway, ca 6 km W of Hilo Roadside. PRESERVED_SPECIMEN Pell, S.K. 830 Hawaii BISH 8/9/2008
50 Toona hexandra (Wall.) M.Roem. East Maui, Hana, along highway 10 mi from town forming overstory of a closed forest with Psidium-Hedychium ferns and grasses as under growth PRESERVED_SPECIMEN Spence, G. 74 Maui BISH 10/15/1972
51 Toona hexandra (Wall.) M.Roem. Lanai flats mauka of pineapple fields; dark forest PRESERVED_SPECIMEN Degener, O. 24561 Lanai BISH 8/13/1957
52 Toona hexandra (Wall.) M.Roem. honolulu: kamanele park Planted. PRESERVED_SPECIMEN degener, o. collector number: 11009 Oahu BISH 5/15/1932
53 Toona hexandra (Wall.) M.Roem. hilo? PRESERVED_SPECIMEN bryan, l.w. collector number: 2 Maui BISH 12/26/1956
54 Toona hexandra (Wall.) M.Roem. Kamanele Park PRESERVED_SPECIMEN Ishikawa, S. 37 Oahu BISH 1/17/1972
55 Toona hexandra (Wall.) M.Roem. honolulu, uh campus PRESERVED_SPECIMEN octavio, a.m. collector number: s.n. Oahu BISH 11/23/1929
56 Toona hexandra (Wall.) M.Roem. Honolulu PRESERVED_SPECIMEN Kerr, M. s.n. Oahu BISH 6/1/1946
57 Toona hexandra (Wall.) M.Roem. e part of hawaii, waiakea arboretum, waiakea forest preserve, 5 mi s of hilo PRESERVED_SPECIMEN e. l. little jr. 23041 BISH 6/2/1968
58 Toona hexandra (Wall.) M.Roem. lanai Mestrosideros-Acacia montane mesic forest 80% closed canopy, 10-12 m tall, with Cheirodendron trigynum, Alphitonia ponderosa, Antidesma platyphylla, Euphoriba atrococca, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena au PRESERVED_SPECIMEN degener, o. desha, a. collector number: 24561 Kauai BISH 8/13/1957
59 Toona hexandra (Wall.) M.Roem. Uppermost Onini Gulch. Mesic forest to mesic shrubland. PRESERVED_SPECIMEN Hughes, G.D. 25 Molokai BISH 3/4/1992
60 Toona hexandra (Wall.) M.Roem. Vicinity of Maunahui PRESERVED_SPECIMEN Oppenheimer, H.L. H110620 Molokai BISH 11/27/2006
61 Toona hexandra (Wall.) M.Roem. Honolulu: Kamanele Park Planted. PRESERVED_SPECIMEN Degener, O. 11009 Oahu BISH 5/15/1932
62 Toona hexandra (Wall.) M.Roem. cultivated near sinclair library, university of hawaii manoa campus PRESERVED_SPECIMEN herbst, d.r. collector number: 636 Oahu BISH 10/7/1967
63 Toona hexandra (Wall.) M.Roem. Honolulu Harbor PRESERVED_SPECIMEN Adee, K.T. Collector Number: s.n. Oahu BISH
64 Toona hexandra (Wall.) M.Roem. none given HUMAN_OBSERVATION Friday, J.B. Hawaii 1/23/2020
65 Toona hexandra (Wall.) M.Roem. Waiakea Timber Management Area HUMAN_OBSERVATION Friday, J.B. Hawaii 3/12/2018
66 Toona hexandra (Wall.) M.Roem. Wilhelmina Rise, HOnolulu PRESERVED_SPECIMEN Mitchell, D.D. s.n. Oahu BISH 10/1/1943
67 Toona hexandra (Wall.) M.Roem. Waipio Valley HUMAN_OBSERVATION Friday, J.B. Hawaii 1/15/2012
68 Toona hexandra (Wall.) M.Roem. Paauilo HUMAN_OBSERVATION Friday, J.B. Hawaii 3/12/2018
69 Toona hexandra (Wall.) M.Roem. honolulu: kamanele park PRESERVED_SPECIMEN degener, o. collector number: 8776 Oahu BISH 5/1/1932
70 Toona hexandra (Wall.) M.Roem. Honolulu, UH Campus PRESERVED_SPECIMEN Octavio, A.M. s.n. Oahu BISH 11/23/1929
71 Toona hexandra (Wall.) M.Roem. waiakea arboretum,waiakea forest preserve,5m,s.of hilo Secondary forest. PRESERVED_SPECIMEN little jr., e.l. collector number: 23041 Hawaii BISH 6/2/1968
72 Toona hexandra (Wall.) M.Roem. honolulu, tantalus On coral reef near shore in shallow water. -- June 1965 PRESERVED_SPECIMEN ewart iii, g.r. collector number: s.n. Oahu BISH 3/31/1929
73 Toona hexandra (Wall.) M.Roem. Hawaii County, Hawaii Island, Roadside on Stainback Highway, ca 6 km W of Hilo Roadside. PRESERVED_SPECIMEN Pell, S.K. Bess, E.; Rannekleiv, K. Collector Number: 830 Hawaii BISH 8/9/2008
74 Toona hexandra (Wall.) M.Roem. honolulu, kamanele park PRESERVED_SPECIMEN degener, o. collector number: 8738 Oahu BISH 8/8/1932
75 Toona hexandra (Wall.) M.Roem. Kalaupapa, E end PRESERVED_SPECIMEN Oppenheimer, H.L. H60727 Molokai BISH 6/30/2007
76 Toona hexandra (Wall.) M.Roem. Honolulu, Tantalus PRESERVED_SPECIMEN Ewart III, G.R. s.n. Oahu BISH 3/31/1929
77 Toona hexandra (Wall.) M.Roem. Honolulu PRESERVED_SPECIMEN Kerr, M. Collector Number: s.n. Oahu BISH 6/1/1946
78 Toona hexandra (Wall.) M.Roem. wailua game reserve, above wailua. begetaion on plots stripmined for bauxite ore in 1958-60. tree plot area. fertilizer applied topically PRESERVED_SPECIMEN howard, r.a. collector number: 20226 Kauai BISH 10/20/1985
79 Toona hexandra (Wall.) M.Roem. wilhelmina rise, honolulu shaded rain forest. with Dryopteris paleacea, Sadleria, Athyrium, Polystichum, Metrosideros, Pelea, Rubus, Styphelia, Cheirodendron, Vaccinium, etc. PRESERVED_SPECIMEN mitchell, d.d. Maui BISH 10/1/1943
80 Toona hexandra (Wall.) M.Roem. hawaii county, hawaii island, roadside on stainback highway, ca 6 km w of hilo lower, dryish forest PRESERVED_SPECIMEN pell, s.k. bess, e.; rannekleiv, k. collector number: 830 Kauai BISH 8/9/2008
81 Toona hexandra (Wall.) M.Roem. honolulu PRESERVED_SPECIMEN kerr, m. collector number: s.n. Oahu BISH 6/1/1946
82 Toona hexandra (Wall.) M.Roem. Honolulu, UH Campus PRESERVED_SPECIMEN Woolford, H.A. s.n. Oahu BISH 3/12/1952
83 Toona hexandra (Wall.) M.Roem. kalaupapa, e end PRESERVED_SPECIMEN oppenheimer, h.l. garnett, b.; czar, g. collector number: h60727 Molokai BISH 6/30/2007